Hygrophorus subgen. Camarophylli (as Camarophyllus) Fr., Summa veg. Scand., Section Post. (Stockholm) 2: 307 (1849). Type species Agaricus camarophyllus Alb. & Schwein., Consp. Fung. Lusat.: 177 (1805) : Fr., [Art. 22.6] ≡ Hygrophorus camarophyllus (Alb. & Schwein. : Fr.) Dumée, Grandjean & L. Maire, Bull. Soc. mycol. Angiogenesis inhibitor Fr. 28: 292 (1912), [= Hygrophorus caprinus (Scop.) Fr. (1838), superfluous to a sanctioned

name, nom. illeg., Art. 13.1]. Hygrophorus subgen. Camarophylli emended here by E. Larss. to exclude A. pratensis and related species now placed in Cuphophyllus. Pileus surface usually dry, gray, grayish blue, buff brown, reddish brown, bistre or fuliginous, or if glutinous then white with yellow floccose-fibrillose veil remnants on the margin; lamellae subdecurrent to decurrent; stipe surface dry, smooth or fibrillose, usually pale gray, grayish blue, buff brown,

bistre or fuliginous, if white glutinous with yellow floccules from veil remnants especially near the apex; lamellar trama divergent giving rise directly JQ1 order to basidia, thus differing from the genus Cuphophyllus. Phylogenetic support Our LSU analysis shows moderately high support (72 % MLBS) for H. chrysodon (subg. Camarophylli) as basal to the rest of the genus Hygrophorus. One ITS analysis (Online Resource 3) shows the same topology while another (Online Resource 9) shows H. chrysodon near the base, both without significant BS support. A four-gene analysis with more species presented by E. Larsson (2010 and unpublished data) also shows subg. Camarophylli as a basal group in Hygrophorus, where it appears as a paraphyletic grade (55 % MPBS for the Resminostat branch separating it from subg. Colorati). Hygrophorus chrysodon and H. camarophyllus appear together in a basal clade in one of our ITS ML analyses (not shown), but H. subviscifer also appears in the clade, and BS support is lacking. Our Supermatrix

analysis places H. chrysodon among sections of subg. Colorati, but without backbone support. Sections included Type section Camarophylli P. Karst., sect. Chrysodontes (Singer) E. Larss., stat. nov. and a new section to accommodate H. inocybiformis, sect. Rimosi E. Larss., sect. nov., are included based on morphology and molecular phylogenies. Comments Agaricus camarophyllus was included by Fries 1821 in his ’subtrib. Camarophylli’ (invalid, Art. 33.9). In 1838, Fries presented this taxon in his’trib. Camarophyllus’ (invalid, Art. 33.9) as Agaricus caprinus Scop., with A. camarophyllus in synonymy. The first valid publication of subgen. Camarophyllus by Fries was in 1849. Fries’ Hygrophorus subg. Camarophylli comprised the type species (H. camarophyllus), H. nemoreus (now placed in Hygrophorus subg. Colorati) and two species of Cuphophyllus (C. pratensis and C. virgineus), so we only retain Fries’ type species.